Jacksonville Jaguars LB Dan Skuta (hip), DE Dante Fowler (shoulder), DE Jared Odrick (triceps) and TE Ben Koyack (knee) did not participate in practice Thursday, Sept. 30. LB Hayes Pullard (ankle) was limited during practice, while OT Kelvin Beachum (concussion), CB Prince Amukamara (hamstring), C Brandon Linder (knee) and DT Malik Jackson (quadriceps) fully participated in practice.